mornin! so my attempt at a terrarium was a miserable fail, so please take this worth less than a grain of salt. but, reading your second post hit on what i was thinking and that is the fan and air circulation. the fans made the biggest difference when we had our us and running. and when one fan died ( we had 2 going in there) things started to go downhill. so, for enclosed space growing my experience was that air flow is critical.
that’s where I would start, but im sure other more experienced tank growers will chime in with better advice. also, it doesn’t look like bugs to me either, it seems more climatic. best of luck!
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